Emergencia de las formas vivas: aspectos dinámicos de la biología

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Emergencia de las formas vivas: aspectos dinámicos de la biología evolutiva Lorena Caballero Universidad Nacional Autónoma de México

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Ciudad de México CopIt ArXives 2008

CopIt ArXives Ciudad de México Copyright © 2008 por Lorena Caballero

;EWLMRKXSR (' Publicado 2008 por CopIt ArXives

Todos los derechos de propiedad de esta publicación pertenece al autor quien, no obstante, concede su autorización al lector para copiar, imprimir y distribuir su trabajjo libremente, en partes o completo, con la única condición que (i) el nombre del autor y y el título original del trabajo sea citado en todo momento, (ii) que el texto no sea modificado o mezclado y (iii) el uso final de este contenido no debe ser comercial, el incumplimiento de estas condiciones será una violación a la ley.

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[email protected] 89 PCF9G @C DI989B <579F 9B [email protected] [email protected] 89 9GD97=9 C 5 8=GH=BHCG [email protected] @9>5BCG [email protected] [email protected] 89 9GD97=9 B 9GH9 HF565>C G9 D5FH9 89 @5 =895 89 9G9B7=5 D5F5 7CB796=F 5 @5 9GD97=9 M [email protected]=75F 5 D5FH=F 89 @5 H9CF\5 89 @CG G=GH9A5G 8=BZA=7CG 89 EI[ A5B9F5 9L=GH9B @5G 9GD97=9G G 897=F DFCDCB;C 5 @5G 9GD97=9G 7CAC 9G9B7=5G ;9B9F585G DCF 5HF57HCF9G 9LHF5]CG ACF:C;9B[H=7CG 7CB @5 75D57=858 89 [email protected]=CB5F BC G^@C 7CAC F9GDI9GH5 8=F97H5 5 @5 577=^B 89 @CG ;9B9G +9 HF5H5 5G\ 89 7CB796=F 5 @5 9GD97=9 7CAC IB G=GH9A5 EI9 BC 9G IB5 [email protected] 7CBGHFI77=^B [email protected] D9BG5A=9BHC C BC H=9B9 @5 J=FHI8 89 [email protected] IB G=GH9A5 89 [email protected]=O757=^B BCJ98CGC EI9 [email protected] [email protected] [email protected] EI9 [email protected] 9B 9GH9 @=6FC 89BHFC 89 @5 [email protected];\5 +=B 9A65F;C @5 8=G7IG=^B EI9 AI9GHFC A9 D5F979 =ADCFH5BH9 [email protected] M D5F5 @CG C6>9H=JCG 89 9GH9 H9LHC BCG =BHFC8I79 89 A5B9F5 6F9J9 IB5 D5FH9 :[email protected] 89 @CG [email protected]=9BHCG 89 @5 [email protected];\5 [email protected]=J5 [email protected] [email protected]@C M @5 [email protected] 89BHFC 89 @5 =BJ9GH=;57=^B [email protected]=B5F=5 9BHF9 @5G 7=9B7=5G 89 @5 [email protected]>=858 M @CG 9GHI8=CG 9JC89JC

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F57=5G 5 @5 =BPI9B7=5 89 D9BG58CF9G 7F\H=7CG 7CAC [email protected] M $9KCBH=B    M [email protected] GI DFCDI9GH5 GC6F9 [email protected] [email protected]=6F=C DIBHI58C 9B 8CB89 F97CBC79B @5 [email protected]=858 89 IB5 [email protected]=^B BC @[email protected] 9AD=9N5 5 F9 [email protected] [email protected] @I;5F [email protected] [email protected]@C 9B @5 7CADF9BG=^B [email protected] DFC79GC [email protected]=JC 9 9GH9 AC8C G9 F97CBC7=^ @5 =ADCFH5B7=5 89 =BHFC8I7=F @CG 7CB79DHCG 89 @5 [email protected];\5 [email protected] [email protected]@C D5F5 7CADF9B89F DFC79GCG BC [email protected] DCF @5 ;9B[H=75 M @5 [email protected]=^B M 89 5<\ @5 B979G=858 89 =B7CFDCF5F 5 @5 ;9B[H=75 [email protected] [email protected]@C M 5 @5 [email protected]=^B 7CAC IB A=GAC [email protected] IB [email protected] 89 HF9G 7I9FDCG 9G 897=F @5 =BH9F577=^B 9BHF9 J5F=CG [email protected] EI9 ;9B9F5B IB G=GH9A5 [email protected]>C J[5G9 CC8K=B  $5 [email protected];\5 [email protected]=J5 [email protected] [email protected]@C 7CAC G9 @9 <5 BCA6F58C 5 9GH5 BI9J5 7CFF=9BH9 89 7CBC7=A=9BHC 9 =BJ9GH=;57=^B GIF;=^ 5 D5FH=F 89 @CG =BH9F9G9G 89 @CG 9A6F=^@C;CG 7CAD5F5H=JCG [email protected] G=;@C 0!0 YGH5 BC 7CAC H9CF\5 :[email protected] H=9B9 5BH97989BH9G 9B @5G =895G 89 F=GH^[email protected] M DCGH9F=CFA9BH9 9B [email protected] G=;@C 0!0 9F5 D5FH9 =ADCFH5BH9 89 @5G =895G 89 J5F=CG =BJ9GH=;58CF9G 7CAC #[email protected] FBGH .CB 59F FBGH [email protected] %[email protected] +9FF9G [email protected]:CIF M 8IF5BH9 [email protected] G=;@C 00 %CF;5B 89 99F /588=B;HCB [email protected]5G IBHC 7CB CHFCG A975B=GACG ;9B9F5B D5HFCB9G 89 [email protected]@C 7CBG9FJ58CG M [email protected] A=GAC H=9ADC AIM 8=:9F9BH9G 9B 9GD97=9G AIM 79F75B5G [email protected] 9B 9GH58=CG AIM H9ADF5BCG [email protected] [email protected]@C  GIF;9 @5 8I85 579F75 89 @5G ;[email protected]=N57=CB9G @[email protected] 9B @5 ;9B[H=75 [email protected] [email protected]@C J[5G9 *5V  GH9 ;F5B 89G7I6F=A=9BHC [email protected]^ 5 @CG =BJ9GH=;58CF9G GC6F9 @C [email protected]=758C EI9 9G <579F ;[email protected]=N57=CB9G 579F75 89 @CG D5HFCB9G 89 [email protected]@C I9 9J= 89BH9 EI9 =65B 89 IB 9LHF9AC 5 CHFC 6IG75B8C @9M9G EI9 9B;@C65F5B HC8C [email protected] DFC79GC <57=9B8C 9LD9F=A9BHCG 75IG5 9:97HC EI9 BC DFC8I7\5B @CG 9:97HCG 9GD9F58CG  D5FH=F 89 9GH5 [email protected] G9 8=9FCB 5 @5 H5F95 89 HF565>5F 7CB



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65G9 9B IB5 =895 8=:9F9BH9 [email protected] [email protected]@C S&C 9G GIO7=9BH9 F9GDCB89F 3DF9;IBH5G 7CB F9GD97HC [email protected] 89G5F [email protected]@C4 8=7=9B8C EI9 G9 HF5H5 [email protected] 89 57H=J5F [email protected];IBCG ;9B9G M 89G57H=J5F CHFCG 9B [email protected] ACA9BHC 7CFF97HC += 6=9B 9G 7=9FHC EI9 @5 [email protected];\5 [email protected]@5F DFCJ99 BIA9FCGCG M [email protected]@58CG DF97989BH9G GC6F9 @CG A975B=GACG DCF @CG [email protected] 9B DF=B7=D=C [email protected]@C G9 [email protected]=N5 F9EI9F=ACG G=B 9A65F;C [email protected];C AZG EI9 9GH5G [email protected]=CB9G 56 [email protected] J9F8589F5G [email protected] J57\5GT +M8B9M F9B9F 9B *C69FH  $5 =B79FH=8IA6F9 579F75 89 @CG A975B=GACG CBHC;[B=7CG M @5 A5B9F5 9B @5 EI9 [GHCG G9 [email protected]=CB5B 7CB @5 [email protected]=^B ;9B9F^ [email protected] B57=A=9BHC 89 9JC89JC 7CAC [email protected] 89 @5 B979G=858 89 9B7CBHF5F D5HFCB9G ;[email protected] 89BHFC 89 @5G F989G [email protected]>5G 89 @5 HF5M97HCF=5 CBHC;9B[H=75 M J9F 89BHFC 89 [GH5G @CG DFC79GCG EI9 D9FA=H9B @5 [email protected]=^B <57=5 CHF5G :CFA5G C [email protected] @5 7CBG9FJ57=^B 89 @5G :CFA5G DF99L=GH9BH9G B @5 [email protected]=858 @5 =ADCFH5B7=5 89 @5 7CCD9F57=^B 9BHF9 @5 [email protected];\5 [email protected] [email protected]@C M @5 [email protected]=^B G9 [email protected] 7CB ;F5B :I9FN5 9B @CG [email protected]=9BHCG 9JC89JC JC 89JC 6IG75 [email protected]=CB5F [email protected] [email protected]@C [email protected] 9A6F=^B 7CB @5 [email protected]=^B M G9 [email protected] @5G G=;I=9BH9G DF9;IBH5G W^AC 5:97H5 [email protected] DFC79GC 89 [email protected]@C 9A6F=CB5F=C 5 @CG 75A6=CG [email protected]=JCG EI9 GI:F9B @CG G9F9G J=JCG M 7^AC [email protected]=CB5 [email protected] [email protected]@C 9B G\

W^AC AC8=O757=CB9G C 75A6=CG ACF:[email protected]^;=7CG =BPIM9B 9B @5 5D5F= 7=^B 89 BI9J5G 9GD97=9G C S&CJ98589G [email protected]=J5GT JC89JC =BH9BH5 85F IB @I;5F :[email protected] 5 @5 9A9F;9B7=5 7CAC IB DFC 79GC =BHF\BG97C [email protected] [email protected]@C [email protected] A=GAC H=9ADC EI9 HF5H5 89 7CADF9B89F @5 [email protected]=7=858 585DH5H=J5 M :9BCH\D=75 EI9 H=9B9 [email protected] 9A6F=^B 9B @5 [email protected]=^B M 89G89 @I9;C @5 =ADCFH5B7=5 89 @5 [email protected]=^B EI9 H=9B9 8=7CF <9FF5A=9B H5 D5F5 9BH9B89F [email protected] AIB8C 89 @5G :CFA5G += 89BHFC 89 9GH9 DFC79GC 9B [email protected]

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[email protected]=^B <5 G=8C GI6CF8=B58C 89 [email protected];IB5 A5B9F5 9B @5 5;9B85 [email protected]^;=75 +=B 9A65F;C GI =ADCFH5B7=5 D9FA5B979 @5H9BH9 M 5`B BC [email protected] HC8C [email protected] B [email protected] A5F7C 89 @5G 8=G7IG=CB9G [email protected] G=;@C 0!0 579F75 89 @5G
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7CB79D7=CB9G 89 GI H=9ADC M BIB75 579DH^ @5G =895G GC6F9 @5 9JC@I7=^B =B7@IGC 89GDI[G 89 @5 DI6@=757=^B 89@ CF=;9B 89 @5G 9GD97=9G 89 5FK=B   -B D9FGCB5>9 AIM =ADCFH5BH9 9B 9@ :CFH5@97=A=9BHC 89 @5 6=C@C;\5 9JC@IH=J5 :I9 <5F@9G 5FK=B (5F5 5FK=B 9@ IGC 89 9A6F=CB9G D5F5 HF5N5F F9@57=CB9G 9JC@IH=J5G 9F5 J=H5@ F;IA9BH565 EI9 9B @CG 9A6F=CB9G 89 5B=A5@9G DC8\5ACG J9F @5 F9@57=^B 9GHFI7HIF5@ 7CB GIG DFC;9B=HCF9G (CF 9>9AD@C @5G 9GHFI7HIF5G @5FJ5@9G BCG D9FA=H9B F9@57=CB5F 7=9FHCG CF;5B=GACG EI9 5 G=AD@9 J=GH5 D5F979B BC H9B9F J\B7I@CG 7@5FCG 7CB 9@ ;FIDC 5@ EI9 D9FH9B979B 5FK=B C6G9FJ^ EI9 9A6F=CB9G 89 5B=A5@9G 89 7I5@EI=9F D=75A9BH9 9@ D9GC EI9 @9 8=C 9B GIG D@5BH95A=9BHCG 5 @5G :I9FN5G 89 G9@977=^B B5HIF5@ =B7@=B^ @5 65@5BN5 D5F5 EI9 89 5@;IB5 A5B 9F5 @5 =ADCFH5B7=5 89@ 9A6F=^B 9B @5 9JC@I7=^B BC :I9F5 H5B =B7@IG=J5 9B @5G H9CF\5G 9JC@IH=J5G AZG 579DH585G 96=8C 5 @5 5D5F=7=^B 89 @5 H9CF\5 89 5FK=B M DCGH9F=CFA9BH9 @5 DFC @=:9F57=^B 89 9GHI8=CG 9B ;9B[H=75 @5 9A6F=C@C;\5 G9 G9D5F^ 89 @CG 9GHI8=CG 9JC@IH=JCG M ;F57=5G 5 @5 =BPI9B7=5 89@ DFC;F5A5 9LD9F=A9BH5@ DFCDI9GHC DCF /=@<9@A *CIL      9B  9@ 9GHI8=C GC6F9 9@ 9A6F=^B G9 79BHF^ =B7@IGC <5GH5 BI9GHFCG 8\5G 9B @5 9LD@=757=^B ;9B[H=75 89@ 89G5FFC@@C $5 =BPI9B7=5 89 /=@<9@A *CIL 9B @5 G9D5F57=^B 89 @5 9A6F=C@C;\5 89 @5G H9CF\5G 9JC@IH=J5G G9 896=^ 5 GIG G^@=85G 65G9G 9LD9F=A9BH5@9G 89 <97
 GH9 <97^ 89 @58C @CG 9GHI8=CG H9^F=7CG EI9 9B 8=:9F9BH9G ACA9BHCG G9 <5 HF5H58C 89 F9G75H5F *CIL D9BG565 EI9 @CG 9A6F=CB9G 9F5B @5 7CBGHFI77=^B 89 IB 89G5FFC@@C 9B ACG5=7C 9B 9@ 7I5@ 7585 D=9N5 89@ CF;5B=GAC 7CBH9B\5 =B:CFA57=^B 8=:9F9BH9 89@ A5H9F=5@ BI7@9=7C ;9B9F5B8C IB5 =895 89 9GD97=5@=N57=^B 9 =BH9;F57=^B 9B IB G^@C 7CB79DHC $5 <=D^H9G=G 89 @5 EI9 D5FH\5 9F5 EI9 G= 9@


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Emergencia de las formas vivas: aspectos dinámicos de la biología

Emergencia de las formas vivas: aspectos dinámicos de la biología evolutiva Lorena Caballero Universidad Nacional Autónoma de México -7&2 Ciudad de ...

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